My immediate reaction, when I saw the photos, was dogs. Maybe coyotes, but coyotes would probably have eaten a lot more. I also wonder how much the actual killer(s) ate. When that happens in this area the dogs basically run the cow to death, or near death, and then run off. Then the scavengers move in. That last case near us was a cow that was run to the point of breaking a leg. There were no marks on the cow. The owner had to put the cow down.