A couple things stand out to me here.
1) Where does science draw the line dividing a reflexive action/ adapted ability/ learned response, and an actual, very basic thought process, where fish are concerned? I've mentioned this here before, but I still struggle to place every observed behaviour under the category of instinct, or adaptive behaviour rather than something more profound?
2) This particular study appears to reinforce a strong belief of my own: adding Tilapia to a traditional, LMB heavy, trophy BG pond may not be in your best interests if continued management of trophy BG is the goal, especially in the presence of vegetation.