So the urogenital opening is the decisive factor? Where does that leave the different colors on the opercular theory standing?

Not doubting you at all. 2 does appear gravid, do the differences in urogenital orifices manifest themselves that vividly at times other than spawn or pre-spawn?

"Forget pounds and ounces, I'm figuring displacement!"

If we accept that: MBG(+)FGSF(=)HBG(F1)
And we surmise that: BG(>)HBG(F1) while GSF(<)HBG(F1)
Would it hold true that: HBG(F1)(+)AM500(x)q.d.(=)1.5lbGRWT?
PB answer: It depends.