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I suspect slow growth if the RES in these ponds as with most slower growing fish was due too many individuals over eating the available food source. This is a very common problem in most ponds. The author did mention that the ponds were at estimated carrying capacity in terms of total fish poundage.
Sounds like from a management standpoint, the ponds needed some heavy angling pressure and/or seining to remove some of the biomass to let remaining fish be more prosperous.
